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The top HVAC controller (has the digital temperature readout) is for the automatic HVAC system. The second one with just the temperature sliders is for the manual HVAC system. To the best of my knowledge, they are not interchangeable.
The automatic HVAC controller relies on several different interior temperature sensors to maintain the passenger compartment at the temperature that is dialed into the system. Let's say you set the temperature for 72° F and the controller is set to automatic. If the temperature outside is 92° F, the HVAC controller commands the AC compressor to turn on, and it also commands the HVAC actuators to send most if not all of the air out of the dash vents. Now, if the outside temperature falls to 40° F, the HVAC controller commands that the temperature HVAC actuators to move to allow heated air from the heater core to blend with the cooled air from the AC evaporator to so that the air coming out of the HVAC vents is at a warmer temperature such that the passenger compartment is maintained at 72° F.
Now, with the manual HVAC controller, you have to choose what temperature you want when you want it. You are the one who has to adjust the temperature sliders in order to stay comfortable. Also, you have to manually select if you want only the heater or only want AC.
Now to complicate matters a little more, before the HVAC controller can do it's job (whether automatically or manually), it has to receive the OK for HVAC command from the Body Control Module (BCM). If the BCM does not see the proper HVAC controller present, then it assumes that the HVAC controller is malfunctioning.
Now, I may be wrong in the end and discover that the two controllers are interchangeable, but I doubt it. But, if I am wrong, I will be the first one to admit it.
Good Luck and enjoy your new ride!!!